Map of the study area. 

 Stationary survey locations: 1, Bozatau district; 2, Kanlykul district; 3, Kungrad district; 4, Muynak district; 5, Chimbay district.  Transect survey locations: 6, Nukus district; 7, Shumanay district; 8, Kegeyli district; 9, Khojeyli district; 10, Beruni district.